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Membranes were sedimented by ultracentrifugation at 150,000 g at 4 C for 1 h, and resuspended in 16 mL of solubilized buffer (1%(w/v) DDM, 150 mM NaCl, 10%(v/v) glycerol, 1 mg mL 1 iodoacetamide, 0.2 mM TCEP, 0.1 protease inhibitor cocktail, 50 mM Tris-HCl pH 8.0)