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In addition, under hypoxic conditions, mitochondria can promote self-digestion and regulate ROS levels through physical interactions with lysosomes, a novel structural phenomenon termed megamitochondria engulfing lysosome (MMEL) [158]

As a master antioxidant, GSH directly scavenges reactive oxygen species and serves as a co-substrate for glutathione peroxidase (GPx), underpinning critical processes including detoxification, DNA synthesis, and immune regulation

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